AB, Kenyon College, 1969. Doctor of Philosophy, Georgetown University, 1980.
Program analyst Occupational Safety and Health Administration, unites states department Labor, Washington, 1976-1977; biologist, Food and Drug Administration, Washington, 1977-1980; research associate, George Washington University Medical Center, Washington, 1980-1982; instructor, Prince Georges Community College, Largo, Maryland., 1982; assistant professor veterinary medicine, Texas Agricultural and Mechanical U., College Station, 1982-1987; associate professor, Virginia/Maryland. Regional College Veterinary Medicine, Virginia Polytechnic Institute and State University, Blacksburg, since 1987.
Staff sergeant United States Air Force, 1970-1974, Vietnam. Member American Association Veterinary Anatomists (correspondent secretary 1988-1990, president 1991-1992), American Veterinary Computer Society, Electron Microscopy Society of America, American Fisheries Society, International Association Aquatic Animal Medicine, American Society of Icthyologists and Herpetologists, Texas Society for Electron Microscopy, New York Academy of Sciences, Sigma Xi.
Married Susan Ann Wolfe, June 28, 1975.