Bachelor of Science in Earth Science, U. Washington, 1972; postgraduate, University of Illinois, Chicago, 1972-1974; Doctor of Philosophy in Biology, University of California at Los Angeles, 1981.
Lecturer, biology California State University, Los Angeles, 1981-1983. Science chair Windward School, 1983—1997, since 2002. Research biologist University of California at Los Angeles, 1985—1998.
Director Windward Conservation Biology Institute, since 1997. Adjunct professor biology California State Polytechnic University, Pomona, since 1991. Consultant United States Army Corps of Engineers, Los Angeles, 1979-1980, California Department Fish and Game, since 1986, Metropolitan Water District, Los Angeles, 1991, 93, 94, 2004, 2005, Department Public Works, Los Angeles County, 1991, 94—, United States Fish Wildlife Service, since 1992, Perrier Corps of America, since 1998.
Chair Southern California Native Fishes Working Group, since 1996.
Member American Association for the Advancement of Science, American Society Ichthyology and Herpetology, American Fisheries Society, Desert Fishes Council, European Ichthyological Congress, Society Conservation Biology.
Married Doris Anne Mendenhall, October 22, 1988. 1 child, Victoria Tamsin.