Master of Science in Biochemistry, Moscow State University, 1973. Doctor of Philosophy in Molecular Biology, Engelhart Institute, Moscow, 1981.
Junior researcher, researcher, senior researcher Russian Academy of Sciences, Puschino, Moscow, 1973-1990. Research associate University Rochester, New York, 1991-1997. Research scientist University Texas, Austin, since 1997.
Member American Society for Biochemistry and Molecular Biology.