Bachelor of Science, Brigham Young University, 1961. Master of Science, University Wisconsin, 1964. Doctor of Philosophy, University Wisconsin, 1967.
Assistant professor, U. Montana, Missoula, 1969-1972; associate professor, U. Montana, Missoula, 1972-1975; professor, U. Montana, Missoula, since 1975. Director Montana Center of Excellence in Biotechnology, Missoula.
Program director National Science Foundation, Washington, 1989-1990. Fellow American Association for the Advancement of Science. Member American Society for Biochemistry, American Society for Microbiology, Biophysics Society.
Married June 8, 1961. Children: Heber, David, Virginia, Kathryn, Kenneth, Elizabeth, Bryce.