Bachelor, Yale University, 1962. Master of Science, Yale University, 1964. Doctor of Philosophy, Yale University, 1967.
From assistant professor to associate professor Stanford (California) University, 1969-1985, professor, since 1985.
Trustee Rocky Mountain Biological Laboratory, Crested Butte, Colorado, 1971-1993, California Academy of Sciences, San Francisco, since 1997. Fellow American Association for the Advancement of Science. Member National Center for Science Education, Society for Study of Evolution, Genetics Society of America, Arctic Institute North America.
Married Carol L. Boggs. Children: Jean Godfrey-June, Laura.