Bachelor of Science, National National Chung-Hsing University University, Taichung, China, 1956. Master of Science, Utah State University, 1964. Doctor of Philosophy, University Utah, 1967.
Postdoctoral fellow Stanford (California) University, 1967-1969. Professor cytogenetics Michigan State University, East Lansing, 1969-1982. Professor genetics University Manitoba, Winnipeg, Canada, 1982-1988.
Curator, project coordinator Missouri Botanical Garden, St. Louis, since 1988. Visiting professor National National Chung-Hsing University University, 1975-1976. Honorary professor Academic Sinica, Beijing, 1980, Jiangsu Agriculture College, 1984.
Consultant World Bank, 1984.
Lieutenant Republic of China Army, 1956-1958. Member Agronomy Society of America, American Society Plant Taxonomists.
Married Nancy Shen, February 16, 1963. Children: Daniel, Dean, Cynthia, Carolyn.