Bachelor of Science, California Institute of Technology, 1959. Doctor of Philosophy, University Washington, 1964.
Postdoctoral fellow, Rockefeller U., New York City, 1964-1965;
research associate, Rockefeller U., 1965-1968;
assistant professor, University of Illinois, Urbana, 1968-1975;
associate professor, University of Southern California, Los Angeles, 1975-1979;
professor biology, professor neurology, University of Southern California, since 1979;
vice president science affairs, Nelson Research & Development Company, Irvine, California, 1981-1982;
acting vice president research & development, Nelson Research & Development Company, 1985-1986;
director program neurological information science, University of Southern California, 1982-1992;
director program in psychobiology, University of Southern California, since 1991. Director cellular biology University of Southern California, 1979-1981, director neurobiology, 1982-1988, director progressive psychobiology, since 1991. Consultant in field; director Marine & Freshwater biomedical Center, University of Southern California, 1982-1983.
Co-director Baja California Expedition of the R/V Alpha Helix, 1974, others. Chairman Winter Conference onBrain Research, 1979, 80, others. Lecturer in field; science advisory board NelsonR & D, 1972-1991.
Member board commentators Brain and Behavioral Sciences, since 1978.
Board directors San Pedro and Peninsula Hospital Foundation, 1989—1995, San Pedro Health Service, 1992—1997. Member of American Association for the Advancement of Science, New York Academy of Sciences, Bay Surgical Society, Association Neurosci. Department and Programs, International Society Neurochemistry, American Society Biological Chemistry and Molecular Biology, Society Neurosci., American Society Neurochemistry, University Park Investment Group, University Club University Southern California (board directors 1991-1995, since 2000), Phi Beta Kappa, Phi Kappa Phi.
Married Pamela Preston Harris, March 9, 1968 (divorced 1979). Children: Heather Harris, Rexford Owen. Married Sara Joan Rorke, July 27, 1980.