Bachelor, Washington University, 1969; Master of Science in Statistics, University of Michigan, 1972; Doctor of Philosophy in Genetics, University of Michigan, 1972.
Junior fellow Michigan Society Fellows, Ann Arbor, 1972-1974. Assistant professor University Texas, Austin, 1974-1977. Associate professor Washington University, St. Louis, 1977-1981, professor, since 1981, Charles Rebstock professor biology, since 2001.
Professor Institute Evolution, Haifa, Israel, since 2007. Consultant St. Louis Zoological Park, since 1979. Founding member, director Society for Conservation Biology, since 1985.
Fellow American Association for the Advancement of Science. Member Society for Study Evolution (vice president 1982, president 1996-1997), Society Conservation Biology (board directors 1985-1988), Nature Conservancy (trustee Missouri chapter since 1988, vice president 1996-2000).
Married Bonnie A. Altman, December 20, 1969. Children: Jeremy Alan, Jeffrey Alan.