Master of Arts with honors, Edinburgh (Scotland) University, 1969. Doctor of Philosophy, Cambridge (England) University, 1973. SKMB certified, Basel (Switzerland) University, 1975.
Postdoctoral fellow biozentrum, Basel U., 1973-1978;
senior investigator, National Institutes of Health, Bethesda, Maryland., 1978-1985;
section chief, National Institutes of Health, Bethesda, Maryland., 1985-1990;
laboratory chief, National Institute Arthritis Musculoskeletal & Skin Diseases, Bethesda, since 1990. Member National Institutes of Health/BRT Study Section, Bethesda, 1987-1992. Chairman Science Technology Engineering And Mathematics advising committee Brookhaven (New York) National Laboratory.
Chairman advising committee on computers in biomedical research, National Institutes of Health, Bethesda, since 1988.
Married Ray Esther Findlay, June 1, 1971. Children: Mairi Monteith Steven, Colin Robert, Andrew James.