Bachelor, Cornell University, 1950. Doctor of Philosophy, California Institute of Technology, 1953.
Fellow, National Instituts of Health 1953-1954, Biochemist 1954-1960. National Science Foundation Fellow, Labs, of F. C. Crick, Cambridge and F. Jacob, Paris 1961. Chief Section of Microbial Genetics, Laboratory of Molecular Biology, National Instituts of Health 1962-1967.
Professor, of Biochemistry, University of California, Berkeley since 1968, Chairman Department, of Biochemistry 1983-1989. Fellow, American Association for the Advancement of Science. Eli Lilly.
Fellow Academy Toxicol. Sciences, American Academy Microbiology, Gerontological Society of America. Member National Academy of Sciences, American Society Biological Chemists, American Society Microbiology (New Brunswick lecturer 1980, Abbott Lifetime Achievement award 2001), Environmental Mutagen Society (award 1977), Genetics Society, American Association Cancer Research, Society Toxicology (Gustavus John Esselen award 1992), American Chemical Society (Eli Lilly award 1964), Royal Swedish Academy of Sciences, American Academy Arts and Sciences.
Married Giovanna Ferro-Luzzi, August 26, 1960. Children: Sofia, Matteo.