Master of Science in Biology, University Leiden, The Netherlands, 1961. Doctor of Philosophy in Molecular Biology, University Leiden, The Netherlands, 1968.
Post doctoral fellow, California Institute Technology, Pasadena, California, 1968-1969; senior research assistant, Department Physiological Chemistry, Leiden, 1970-1974; associate professor, Department Medical Biochemistry, Leiden, 1974-1980; professor laboratory of molecular carcinogenesis, State University Leiden, Leiden, since 1980. Member recombinant DNA advisory commission The Netherland government, Den Haag, since 1975.
2nd lieutenant Medical Troops, The Netherlands, 1962-1963. Member The royal Netherlands Academy of Sciences and Letters (Beijerinck Virology medal 1978), Holland Science Association (AKZO prize 1975), European Molecular Biology Orng.
Married Titia Aukje Brongersma, July 26, 1961. Children: Jeroen, Taco, Marjolyn.