Bachelor, Cornell University, 1972. Doctor of Philosophy, Stanford University, 1977.
Research associate The Wistar Institute, Philadelphia, 1981-1984, assistant professor, 1985-1988, University Pennsylvania School Medicine, Philadelphia, 1986-1988. Senior scientist Worcester Foundation for Experimental Biology, Shrewsbury, Massachusetts, 1988-1997. Associate professor University Massachusetts, Worcester, 1989-1997, professor Medical Center, since 1997.
Member study section American Cancer Society, Atlanta, since 1995. Member Shrewsbury Committee on Hazardous Substances, Shrewsbury, 1992-1993. Member American Chemical Society, American Society Biochemistry and Molecular Biology, Society for Neurosci., American Society for Cell Biology.
Married Elizabeth J. Luna, June 24, 1974.