Doctor of Medicine, University Córdoba, 1953. Doctor of Philosophy, University Córdoba, 1965.
Instructor Institute Physiology, National U. Córdoba, 1953-1955; instructor department biochemistry, National U. Córdoba, 1956-1958; investigator department biochemistry, National U. Córdoba, 1958-1962; from assistant professor to professor department biochemistry, National U. Córdoba, 1959-1971; professor, department chairman biochemistry, National U. Córdoba, 1971-1985; plenary professor department biochemistry, National U. Córdoba, 1985-1996; professor emeritus, National U. Córdoba, since 1996. Research associate Johns Hopkins University School Medicine, Baltimore, 1961-1965. Career investigator, National Council Science Research, Argentina, since 1972.
Member National Academy of Sciences (vice president 1992-1996, 96-2000, since 2000), National Academy Physical and Natural Sciences, Academy Medical Sciences Cordoba (emeritus), Argentine Society Bio-chemical Investigation (president 1976-1978) Society Biology Córdoba (president 1971-1973), Argentine Society Protozoology (president 1986-1990), Argentine Society Genetics, Bio-chemical Society United Kingdom, Society Protozoologists United States, Entomological Society of America.
Married Beatriz Romagna, March 28, 1958 (divorced June 1978). Children: Víctor Gustavo, Sergio Darío, Antonio, Beatriz. Married Cristina Noemí Gardenal, December 8, 1980.
1 child, Julieta.