Doctor of Medicine, Krakow University, 1957. Doctor of Science, Jagiellonian University, Krakow, 1968. Doctor of Philosophy (honorary), Cleveland State University, 1990.
Doctor of Philosophy (honorary), University Hartford, 1995. Doctor of Philosophy (honorary), State University of New York, Buffalo, 1998.
Assistant Medical Academy, Krakow, 1957-1968. Associate professor Jagiellonian University, 1969-1976, professor, since 1976, director Institute of Molecular Biology, 1977-1981, prorector, 1984-1987, rector, 1987-1990, 1993-1999, retired, 2005.
Member Bio-chemical Society United Kingdom, Polish Bio-chemical Society (chairman Krakow branch 1972-1975), Polish Academy of Sciences, Academy of Sciences & Lettres (Krakow).
Married Anna Marchlewska. 1 child, Justina.