Bachelor of Science in Biology, Rutgers University, 1949; Doctor of Philosophy, Rutgers University, 1952.
Associate microbiologist Ortho Research Foundation, Raritan, New Jersey, 1952-1954. Instructor microbial biochemistry Waksman Institute Microbiology, Rutgers University, 1954-1956. Senior assistant scientist to senior scientist Resident Advisor Taft Sanitary Engineering Center, United States Public Health Service, Cincinnati, 1956-1959.
From associate professor to professor bacteriology University Cincinnati, 1959-1971, head department biological science, 1964-1966. Professor biology University Connecticut, Storrs, 1971—1995, head microbiology section, 1974-1985, dean College Liberal Arts and Sciences, 1992—1995, dean emeritus, since 1995. Senior visiting fellow department biochemistry University Leicester, England, 1967-1968.
Visiting fellow in biochemistry Clare Hall, University Cambridge, England, 1979. Program director for cell biology National Science Foundation, Washington, 1984-1985. Visiting scholar department biology University California San Diego, La Jolla, 1989.
United States representative North Atlantic Treaty Organization Advisory Panel on Collaborative Research Grants, 1990-1992.
Member American Association for the Advancement of Science, American Society for Microbiology.
Son of Antonio and Maria R. Married Marjorie J. Backus, 1953. Children: Stephen, James, Charles.