Doctor of Medicine, Hebrew University Hadassah Medical School, Jerusalem, 1965. Doctor of Philosophy, Hebrew University Hadassah Medical School, Jerusalem, 1969.
Postdoctoral fellow University California, San Francisco, 1969—1972. Associate professor Technion-Israel Institute of Technology, Haifa, 1972—1980, professor, since 1980, distinguished professor biochemistry, Rappaport Faculty Medicine, since 1998. Visiting researcher Fox Chase Cancer Center, Philadelphia, 1978—1981.
With Israel Defense Forces, 1965-1967.
Member of National Academy of Sciences (foreign associate), American Academy Arts & Sciences (foreign honorary), European Molecular Biology Organization.
Married Judith Leibowitz, 1963. Children: Daniel, Yair, Oded.