Bachelor of Science, University British Columbia, Vancouver, 1989. Master of Science, University Toronto, Ontario, Canada, 1991. Doctor of Philosophy, Yale University, New Haven, 1995.
Assistant professor State University of New York, Stony Brook, 1998—2000. Associate professor University Alberta, Edmonton, Canada, 2000—2004. Professor University California, Davis, since 2004, associate director bioinformatics Genome center, 2006—2008.
He is an indifferentist, not caring whether God exists.
Married Kimberly Anne Rannala, August 10, 1988.