Bachelor of Science, Michigan State University, East Lansing, 1961; Doctor of Philosophy, University of California-Berkeley, 1966.
Research associate, J.W. Goethe U., Frankfurt/Main, Federal Republic Germany, 1966-1968; research associate, Rockefeller U., New York City, 1968-1970; faculty, U. Iowa, Iowa City, since 1970; professor biochemistry, U. Iowa, Iowa City, since 1979.
Member American Society for Biochemistry and Molecular Biology, American Chemical Society, Sigma Xi.
Married Rosemary Kuhn, June 13, 1962. Children— Brendan Bryce, Laurel Andrea.