He was born at Gilmore, Saint Charles Company, Missouri, and graduated (Doctor of Medicine) from the University of Missouri in 1901 and (Doctor of Philosophy) from the University of Chicago in 1908. Taught physiology while engaged in advanced studies, and was professor of physiology and pharmacology at Washington University in 1906–1909 and at the University of Pittsburgh after 1909.
University of Missouri. University of Chicago.
He was author of Blood-Vessel Surgery and its Applications (1912) and of contributions on blood reactions and alterations, resuscitation, cerebral and other anæmias, isolated and ungrafted tissues, and sutures and anastomosis of blood vessels. A 2001 book by Hugh East. Stephenson, Junior. and Robert South. Kimpton, America"s First Nobel Prize in Medicine or Physiology:The Story of Guthrie and Carrel, argues that the primary cr for this work should have gone to Guthrie rather than Carrel. However, Guthrie"s head transplant experiments likely prevented his Nobel Prize candidacy status.
