Bachelor of Science, Nottingham (England) University, 1958. Doctor of Philosophy, Bristol (England) University, 1961.
Research fellow Cornell University, Ithaca, New York, 1961-1963. Lecturer, professor Glasgow (Scotland) University, 1963-1994. Director Institute biomedical and Life Sciences, University Glasgow, since 1994.
Publications manager Federation of European Microbiol. Societies, since 1991.
Fellow Royal Society Edinburgh, Institute Biology., Royal Society Arts.
Married Margaret Christina Moir, October 4, 1965. Children: Margaret Claire, Catriona Annual.