Bachelor of Science, University of Chicago, 1960; Doctor of Philosophy, University of Chicago, 1965.
Assistant professor, University of California, Santa Barbara, 1966-1969; associate professor, professor biology, University of Illinois, Chicago, 1969-1995; professor emeritus, University of Illinois, Chicago, since 1995. Visiting professor University College, Dublin, Ireland, 1979-1980.
Fellow American Association for the Advancement of Science. Member Ecological Society American, Society Study Evolution. M C.
Married Sarah Burnham, January 1961 (divorced 1986).