Bachelor of Science in Chemistry, Massachusetts Institute of Technology, 1972. Doctor of Philosophy in Molecular Biology, University of California at Los Angeles, 1977.
Postdoctoral associate, Pasteur Institute, Paris, 1977-1979; postdoctoral associate, Cornell Univercity, Ithaca, New York, 1979-1981; professor biochemistry and molecular biology, Pennsylvania State University, University Park, since 1981; Ernest C. Pollard professor biotechnology, Pennsylvania State University, University Park, 1992. Visiting professor Swiss Federal Technology U., Zurich, 1989-1990, Australian National U., Canberra, 1996-1997.
Member American Association for the Advancement of Science, American Academy Microbiology, American Society Microbiology, American Society Plant Physiology, American Society Biochemistry and Molecular Biology, International Society Plant Molecular Biology, International Society Photosynthesis Research.
Married Rita Jean Corio, July 20, 1974 (divorced August 1988). Married Veronica Lynne Stirewalt, August 24, 1991.