Bachelor of Science in Biology, Morvian College, 1978. Master of Science in Biology, Temple College, 1981. Doctor of Philosophy, Temple University, 1983.
Postdoctoral fellow Moleular Viral Biology Section National Institute for Allergy and Infectious Diseases-LID National Institutes of Health, Bethesda, Maryland, 1983-1986. Research associate Department of Microbiology and Immunology Stanford University/The Palo Alto (California) Veterans Administration Medical Center, 1986-1991, Department of Microbiology and Immunology and GI State University of New York/Northport (New York ) Veterans Administration Medical Center, Stony Brook, since 1991. Consultant molecular and cell biology core facility Stanford (California) University, 1988-1991.
Member American Association for the Advancement of Science, American Society for Microbiology, American Society Virology.
Married Joanne Marie Johnson, May 23, 1981. 1 child, Natalie Anne.