Bachelor of Science, University South Dakota, 1972. Doctor of Philosophy, University South Dakota, 1976.
Undergraduate research assistant biochemistry department University South Dakota, 1969—1972, graduate research assistant, 1972—1976, teaching assistant biochemistry, 1973—1974, postdoctoral research associate, 1976, National Institutes of Health trainee, 1972—1973. Postdoctoral research associate Purdue University, West Lafayette, Indiana, 1976—1978. Postdoctoral fellow Laboratory Chemical Biology, National Institute Arthritis, Metabolism and Digestive Diseases, National Institutes of Health, Bethesda, Maryland, 1978—1980.
Cancer research scientist II Department Molecular Biology Roswell Park Memorial Institute, Buffalo, 1980—1981. Assistant scientist department medicine Mount Sinai Medical Center, Milwaukee, from 1981. Assistant professor University Wisconsin Medical School, Milwaukee, 1981—1985, associate professor, 1985—1986.
Senior biochemist Abbott Laboratories, Abbott Park, Illinois, 1986—1988. Associate professor Rush-Presbyterian-St. Luke's Medical Center, Chicago, from 1988.
Member of American Association for the Advancement of Science, American Heart Association, American Society Biological Chemists, American Chemical Society, Sigma Xi.