Bachelor of Science, Colorado State University, 1959; Doctor of Veterinary Medicine, Colorado State University, 1962; Master of Science, Purdue University, 1967; Doctor of Philosophy, University Wisconsin-Madison, 1971.
Instructor, then assistant professor, Purdue University, West Lafayette, Indiana, 1962-1969; assistant professor, then associate professor, University of Wisconsin -Madison, 1971-1977; professor, University of Wisconsin -Madison, since 1977; professor department Biosciences, University of Wisconsin -Madison, since 1980. Visiting professor University of California-San Francisco, 1977-1978. Member respiratory and applied physiological study section National Institutes of Health, 1988-1992.
Member American Society Veterinary Physiologists and Pharmacologists (president 1982-1984), American Physiological Society, American Veterinary Medical Association, Wisconsin Association biomedical Reserve Education (president 1998-2000).
Married Sharon Kay Cummings, September 9, 1961. Children– Jennifer, Kristine, Bradley.