Bachelor of Science, California Institute Technology, 1957; Master of Science, California Institute Technology, 1958; Doctor of Science, Massachusetts Institute of Technology, 1963.
Research engineer, Dow Chemical Company, Williamsburg, Virginia, 1958-1960; assistant professor, California Institute Technology, Pasadena, 1964-1968; associate professor, Montana State University, Bozeman, 1969-1976; professor, Montana State University, Bozeman, 1976-1978; professor, U. Rochester, New York, 1978-1998; research professor, Montana State University, Bozeman, since 1998.
Served with United States Army, 1954-1955, Japan. Fellow American Association for the Advancement of Science. Member biomedical Engineering Society, Microcirculatory Society, American Institute Medical and Biological Engineering, Society Rheology, Northern American Society Biorheology, International Society Biorheology (president), European Microcirculation Society.
Married Sarah Drew, June 15, 1963. Children— Becky, Bradford.