University of Illinois, University of California at Berkeley, Johns Hopkins University School of Medicine, Baltimore, Maryland.
Intern Barnes Hospital, St. Louis, 1956-1957. Resident in medicine Henry Ford Hospital, Detroit, 1959-1962. United States Public Health Service fellow department human genetics University Michigan, Ann Arbor, 1962-1964, research associate, 1964-1967.
Assistant professor molecular biology and genetics Johns Hopkins University School Medicine, Baltimore, 1967-1969, associate professor, 1969-1973, professor, 1973—1998. Scientist Celera Genomics Corporation, Rockville, 1998—2002. Scientific director synthetic biology & bioenergy J. Craig Venter Institute, since 2002.
Research associate Research Institute Molecular Pathology, Vienna, 1990—1991.
Member of National Academy of Sciences, American Association for the Advancement of Science, American Society Biological Chemists, American Society Microbiology.
Married Elizabeth Anne Bolton, May 25, 1957. Children: Joel, Barry, Dirk, Bryan, Kirsten.