Bachelor, Washington University, St. Louis, 1976; Doctor of Philosophy, University of Wisconsin, 1980.
Postdoctoral fellow, U. Basel, Switzerland, 1980-1983; professor, U. Basel, Switzerland, since 1988; research scientist, ISREC, Lausanne, Switzerland, 1983-1988. Consultant Ciba Geigy AG, Basel, since 1988.
Member American Society Cell Biology, Swiss Bio-chemical Society.
Married Marie-Francoise Le Guern, November 23, 1975. 1 child, Yannick Samuel.