Student, University Liverpool, England, 1984. Diploma, University Freiburg, 1985. Doctor of Philosophy, University Freiburg, 1988.
Postgraduate, University Zurich, Switzerland, 1988.
Group leader University Freiburg, 1988-1992. Postdoctoral fellow Osaka (Japan) Bioscience Institute, 1992-1994. Scientist University Heidelberg, Germany, 1994-1998, University Ulm, since 1996.
Consultant Gödecke Ag, Freiburg, 1990-1992.
Member New York Academy of Sciences.