Bachelor of Science, U. Edinburgh, Scotland, 1959; Bachelor of Medicine, Bachelor of Surgery, U. Edinburgh, Scotland, 1962; Master of Arts by special resolution, University of Oxford, England, 1973.
Honorary lecturer University Edinburgh, Scotland, 1966-1969. Anglo-French research scholar University Paris, 1969. Research officer University Oxford, Oxford, England, 1970-1979.
Fellow and tutor in medicine St. Edmund Hall University Oxford, 1973-1979. Professor zoology University of Hull, England, 1979-1987. Professor neurophysiology University Edinburgh, Scotland, 1987—2003.
Emeritus fellow St. Edmund Hall, University Oxford, England, since 1979. Member District Health Authority Scarborough, England, 1981.
Fellow Royal College Physicians Edinburgh (honorary library since 2001). Member Royal College Physicians London, Physiological Society.
Married J. Patricia Maule, July 18, 1961. 1 child, David.