Doctor of Medicine, University Helsinki, 1981. Doctor of Philosophy, University Helsinki, 1984.
Research and teaching associate University Helsinki, 1981—1986. Postdoctoral research fellow Memorial Sloan-Kettering Cancer Center, New York City, 1986—1989. From senior research scientist to research professor National Public Health Institute, Helsinki, 1989—2005, research professor, since 2005.
Consultant virologist United Laboratories Ltd., Helsinki, since 1982. Junior officer Army, 1984-1985.
Member of International Society for Interferon and Cytokine Research.
Married Anni-Riitta Virolainen, February 13, 1960. Children: Ora I.A., Lina I.A., Ansa P.A., Otto U.A.