Bachelor of Arts in Mathematics, Princeton University, 1972. Doctor of Philosophy in Biochemistry, Princeton University, 1977.
Postdoctoral fellow, Princeton (New Jersey) U., 1976-1977; postdoctoral fellow, Medical Research Council Laboratory Molecular Biology, Cambridge, England, 1977-1981; postdoctoral fellow, University of Michigan, Ann Arbor, 1982; assistant professor, St. Olaf College, Northfield, Minnesota, 1982-1983; from assistant professor to associate professor, University of Wisconsin, Oshkosh, 1984-1995; professor of chemistry, University of Wisconsin, Oshkosh, since 1995. Visiting researcher Cambridge U., 1993.
Member American Society Cell Biology, Sigma Xi.
Married Verena Winterfeld, August 21, 1982. Children: Bjorn Rolf, Tanya Ruth.