Bachelor of Science with honors, Aston University, United Kingdom, 1980. Doctor of Philosophy, King's College, London University, 1984.
Research fellow Cornell University, Ithaca, New York, 1984-1987. Senior research fellow Medical Research Council Center, Cambridge, England, 1987-1991. Lecturer Birmingham (England) University, 1991-1994.
Senior lecturer Bristol University, 1994-1995, reader, 1995-2000, professor, since 2000. Visiting fellow Kyoto (Japan) University, 1985.
Member British Pharmacology Society, Bio-chemical Society.
Married Kathrine Jean Garvey, September 15, 1984. Children: Samuel Edward, Benjamin Seth, Sophie Annabelle.