Bachelor of Science in Genetics, University Birmingham, England, 1952. Doctor of Philosophy, University Birmingham, England, 1956.
Assistant lecturer in genetics, U. Glasgow, Scotland, 1954-1956;
science officer, Animal Virus Research Institute, Pirbright, England, 1956-1960;
National Foundation fellow, California Institute Technology, Pasadena, 1960-1961;
senior science staff experimental virus research unit, Medical Research Council, Glasgow, 1961-1968;
honorary director virology unit, Medical Research Council, Glasgow, 1968-1994;
professor, chair, director, Institute Virology, U. Glasgow, 1968-1994;
professor emeritus, honorary senior research fellow, U. Glasgow, since 1994. Visiting professor National Institutes of Health, Bethesda, Maryland., 1967, Uniformed Superior vena cava syndrome U. Health Sciences, Bethesda, 1985. Visiting fellow Clare Hall, Cambridge, 1986-1987.
Member science advisory group Equine Virology Research Foundation, Newmarket, England, since 1987. Consultant in field.
Served British Army, 1944-1947, Middle East. Fellow Royal Society Edinburgh (Scotland), Institute Biology. Member Genetical Society (trustee since 1971), European Molecular Biology Organization (chair course and workshops committee 1976-1978), Society General Microbiology, Athenaeum.
Married Barbara Naomi Morris, August 22, 1953. Children: Robert John, Anne Barbara Morton, Ian David.