Master of Science, University Helsinki, 1956. Doctor of Philosophy, University Oulu, Finland, 1974.
Senior teacher Kauhava (Finland) Secondary School, 1959-1961. Keeper Perämeri Research Station University Oulu, 1961-1965, assistant department zoology, 1966-1973, manager Perämeri Research Station Hailuoto and Oulu, 1974-1982. Associate professor fisheries, department biology University Jyväskylä, Finland, 1982-1997, head department Finland, 1988-1990, emeritus professor Finland, since 2000.
Member Marine Biology Collegium for Northern Countries, 1982-1984. Member, chairman aquaculture working group Mittnordenkommitte, 1985-1988. Lieutenant (in Reserve) Finnish Army.
Lieutenant Finnish Army Reserve. Member Academy Finland (environmental subcommittee 1980-1982, research grantee 1991-1992), International Limnology Society.
Married Elsa Tellervo Punkeri, April 10, 1966. Children: Tapio, Katariina, Tero, Kosti, Maria, Anna.