Bachelor, University Oulu, 1970. Doctor of Medicine, University Oulu, 1975. Doctor of Philosophy, University Oulu, 1977.
Visiting associate National Institutes of Health, Bethesda, Maryland, 1978-1979. From resident to professor biochemistry University Oulu, Finland, 1980-1995. Professor medical chemistry Karolinska Institute, Stockholm, since 1994.
Associate professor biochemistry, pathology Rutgers Medical School, Piscataway, New Jersey, 1983-1985. Chairman department biochemistry University Oulu, 1987-1995, deputy director bioctr., 1987-1989, scientific director, 1990-1995. Research professor Finland's Cancer Center, 1989-1994.
Member Nobel Committee for Physiology and Medicine, 1996.
Married Marja-Terttu Kukkonen, January 6, 1972. Children: Karl Kristian, Sam Henrik, Jan Erik.