Doctor of Medicine, Eberhard-Karls University Tubingen, Germany, 1980.
Intern, Katharinenhopital, Stuttgart, Germany resident, German Heart Center resident, Ludolf-Krehl-Klinik, Heidelberg, Germany fellow in hypertension, University of Michigan, Ann Arbor, 1981-1982; assistant, German Heart Center, Munich, 1983-1984; assistant, U. Heidelberg, 1984-1992; oberarzt, Humboldt University of Berlin, 1992-1994; group leader, Max Delbrück Center for Molecular Medicine, Berlin, since 1994.
Served as physician Bundeswehr (German military), 1982-1983. Member Dt. Gesellschaft Herz-und Kreislaufforschung.
Married Regina Merkle, August 4, 1984. Children: Johannes, Richard.