Bachelor, Miami University, Ohio, 1956; Master of Arts, Columbia University, 1957; Doctor of Philosophy, Columbia University, 1961.
Boese fellow, Columbia University, New York City, 1961-1962; North Atlantic Treaty Organization and National Institutes of Health fellow, University College London, 1962-1963; research fellow, American Museum Natural History, New York City, 1963-1966; assistant professor anatomy, University of Chicago, 1967-1971; associate professor evolutionary biology & conceptual foundations science, University of Chicago, 1971-1973; associate professor biology & conceptual foundations of science, University of Chicago, 1973-1976; professor biology and conceptual foundations of science, University of Chicago, 1976-1988; professor ecology, evolution, conceptual foundations science, University of Chicago, since 1988; research associate department geology, Field Museum, Chicago, since 1971.
National advisory board Voice of Reason, New York, since 1981. Member American Association of University Professors (president University of Chicago chapter), Society Study Evolution (vice president 1973, 80), American Society Naturalists (vice president 1974-1975), Paleontological Society (councillor 1980-1982), International Society Cryptozoology (board directors), Ecological Society American.
Married Phebe May Hoff, 1959. Children: Katrina, Diana. Married Virginia C. Maiorana, 1974.
Married Towako Matano, 1998.