Master of Science in Biology & Mathematics, University Heidelberg, 1977. Doctor of Philosophy in Biophysics, University Heidelberg, 1981.
Research group leader Goedecke-Parke Davis Pharmaceutical Company, Freiburg, Germany, 1983—1984. Assistant professor pharmacology University Marburg, Germany, 1984—1986. Head research group Max-Planck-Society, Germany, since 1986.
Head research group enzyme biochemistry Max-Planck-Institute Biophysical Chemistry, Goettingen, Germany, since 1987.