Bachelor of Science in Education, Biology, Indiana University of Pennsylvania, 1973. Doctor of Philosophy in Environmental Studies, Duke University, 1978.
Nongame biologist United States Fish and Wildlife Service, Washington, 1980-1985. Senior ecolog. advisor United States Fish and Wildlife Service, seconded to Agency for International Development, 1985-1987, supervisor cooperative research unit, 1988-1991. Vice president resources, planning and economics The Wilderness Society, 1991-1994.
Vice president, director heritage network operations The Nature Conservancy, since 1994. Member board governors Society for Conservation Biology, 1985-1987.
Member American Association for the Advancement of Science, Society Conservation Biology, The Wildlife Society.
Married Elizabeth June Becker, May 2, 1998.