Doctor of Philosophy, Biology, University of Bristol, 1973—1976. Bachelor, Biology, University of Stirling, 1970—1973.
Principal science officer Dept of Agriculture & Rural Development, Belfast, United Kingdom, since 1982. Biologist British Antarctic Survey, Cambridge, United Kingdom, 1977—1982. Lecturer Queen's University Belfast, Belfast, Northern Ireland, United Kingdom, since 1982.
Visiting professor China Agricultural University, Beijing, since 1998.
Married Susan Janet Neil, March 27, 1984.