Bachelor of Science with honors, University London, 1968. Doctor of Philosophy, Exeter University, England, 1971.
Lecturer Makerere University, Kampala, Uganda, 1971—1973. Research fellow Atomic Energy Canada, Pinawa, 1973—1975. Senior lecturer Rhodes University, Grahamstown, South Africa, 1976—1981.
Professor, chair zoology University Natal, Pietermaritzburg, South Africa, since 1981, dean science South Africa, 1986—1989. Director Research Center for African Parrot Conservation, Pietermaritzburg. Lecturer in field.
Fellow: Linnean Society, Zoological Society London, University Natal Fellows.
Married Megan Jane Jones, September 21, 1969. Children: Benjamin John, Ian Robert. Married Sharynne Joy Penwarden, June 17, 1999 (divorced 2001).