Bachelor, Union College, Lincoln, Nebraska, 1965. Master of Science, University Missouri, 1968. Doctor of Philosophy, University Nairobi, Kenya, 1972.
Medical entomologist, Ministry of Health, Nairobi, 1968-1973; senior lecturer, U. Nairobi, 1973-1979; principal research scientist, ICIPE, Nairobi, 1979-1986; deputy director, ICIPE, Nairobi, 1986-1987; senior principal research scientist, ICIPE, Nairobi, 1986-1994; professor zoology, Jomo Kenyatta U., Nairobi, 1995. Vice-chancellor U. Eastern Africa, Baraton, Kenya, since 1996. Commissioner Common for Higher Education, Nairobi, since 1996.
Fellow African Academy of Sciences, New York Academy of Sciences, Kenya National Academy.
Married Gloria Jean Friday, February 4, 1968. Children: Muthoka, Nzisa, Mueni.