Bachelor of Science in Biology, American University Beirut, 1990. Master of Science in Biomedical Engineering, Bogazici University, Istanbul, Turkey, 1993. Doctor of Philosophy in Molecular biology & genetics, Bogazici University, Istanbul, Turkey, 1999.
Research scientist Max Planck Institute, Berlin, 1999—2000. Research group leader University Bonn, Germany, since 2000.
Member of Epigenetic Society, Room Engineers Tripoli.