Career
From 1897 to 1927 he was professor of Experimental Physiology at the University of Utrecht. In addition to his work on the sense of smell, he also conducted research on the human heart. He found that salts of potassium and other radioactive elements stimulated the heart.
His major work was "Die Physiologie des Geruchs" (Physiologie of Olfaction), it appeared in 1895.
An important contribution of Zwaardemaker"s studies is the concept of odor conjugates. Zwaardemaker discovered that certain odors could be prevented from detection by smell senses when mixed with various essential oils.
These combination of odors are referred to as, (or Z-pairs).