Education
He studied medicine at the Universities of Bern and Würzburg, obtaining his doctorate at Bern in 1872.
anatomist pathologist university professor
He studied medicine at the Universities of Bern and Würzburg, obtaining his doctorate at Bern in 1872.
In 1881 he was appointed professor of pathology and director of the pathological institute in Zürich. During the following year he relocated as a professor to the University of Tübingen, and from 1889 to 1905, he was a professor in Freiburg.