Bachelor of Science, Yale University, 1961; Master of Arts (honorary), Yale University Doctor of Philosophy, Harvard University, 1966.
Postdoctoral fellow, U. Geneva, 1966-1967; postdoctoral fellow, Medical Research Council Laboratory of Molecular Biology, Cambridge, England, 1967-1969; assistant professor, then associate professor department molecular biophysics, Yale University, New Haven, 1969-1976; associate professor department of chemistry, Yale University, New Haven, 1976-1979; professor, Yale University, New Haven, since 1979; department chairman chemistry, Yale University, New Haven, 1987-1990.
Fellow American Association for the Advancement of Science. Member American Chemical Society, American Society Biological Chemists and Molecular Biologists, National Academy of Sciences, American Academy Arts and Sciences, Biophysics Society (editor Biophysics Journal 1997-2002, president since 2010).
Married Margaret Sue Murphy, January 30, 1966. Children: Catherine, Philip.