Bachelor of Science, West Liberty State College, 1964. Doctor of Philosophy in Pharmacology, West Virginia University, 1967.
From assistant to associate professor Washington University Medical School, St. Louis, 1971-1981, professor physiology, from since 1982, head department, since 1984, now Edward Mallinckrodt Junior professor cell biology and physiology. Fellow Space Science Research Center, University Missouri, 1967. Arthritis Foundation fellow molecular biology, Vanderbilt University, 1968-1970.
Member British Bio-chemical Society, American Chemical Society, American Physiological Society, American Society Biological Chemists.
Married; 3 children.