Bachelor of Science, Washington College, Chestertown, Maryland., 1942; Doctor of Philosophy, Harvard University, 1950; Doctor of Science (honorary), Washington College, 1982.
Chemist Atlas Powder Company, 1942-1943. Instructor chemistry Northeast Missouri State Teachers College, Kirksville, 1943-1944. Assistant biochemist Massachusetts General Hospital, Boston, 1949-1950.
Member faculty Washington University Medical School, St. Louis, 1950-1962. Professor biochemistry, chairman department Chicago Medical School, 1962-1966. Professor, chairman department physiology Rutgers Medical School, New Brunswick, New Jersey, 1966-1986.
Member biochemistry test committee National Board Medical Examiners. Member sponsoring committee International Conference on Biological Membranes.
Served with United States Naval Reserve, 1944-1946. Fellow American Institute Chemists, New York Academy of Sciences. Member American Society Biological Chemists, American Society Cell Biology, Corporation Marine Biological Laboratory, American Chemical Society, American Association for the Advancement of Science, American Gastroenterological Association, Biophysics Society, Society General Physiologists, American Physiological Society (chairman public affairs committee), Federation American Society Experimental Biology (chairman public information committee), International Union Physiological Sciences (science program committee), Sigma Xi.
Married Mildred Ellen Price, July 19, 1941 (divorced April 1962). Children— Barbara Joan, JonathanTownley. Married Laura Jane Scott, April 13, 1972.