Bachelor of Science in Physics, California Institute of Technology, 1959. Doctor of Philosophy in Biophysics, University California, Berkeley, 1964.
Postdoctoral fellow University Cologne (Germany), 1964-1965, Pasteur Institute, Paris, 1965-1966, Cold Spring Harbor (New York ) Laboratory, 1966-1968. Assistant professor biochemistry Duke University Medical Center, Durham, North Carolina, 1968-1975. Associate professor biology University Missouri, Kansas City, 1975-1977.
Program director genetic biology National Science Foundation, Washington, 1977—2001. Retired, 2001; senior scientist office assistant director biology, behavioral science, 1981-1982. Visiting professor Johns Hopkins University Medical School, Baltimore, 1987—1988.
Chair National Aeronautics and Space Administration Proposal Review Panel for Microbiology, since 2002. Lecturer Sonoma State University, Rohnert Park, California, 2002. Senior associate Centers Development Initiative, since 2003.
Legislation assistant Congressman Dave McCurdy of Oklahoma, 1980-1981. 1st lieutenant United States Air Force Reserve, since 1962. Fellow American Association for the Advancement of Science.
Member Genetics Society of America, American Society Microbiology.
Married Jenny Elizabeth Flack, June 12, 1959. 1 child, Marc Stuart.